This research present the results of detailed systematic study about the megaspores found in the sedimentary rocks of Itararé Subgroup (Upper Carboniferous) located at the northeastern portion of Paraná Basin (São Paulo State), Brazil. The studied outcrop was deposited by subaquosos fans located at the lower portion of Itararé Subgroup with glacial influence during the Westaphlian times. Four species of megaspores are here described and illustrated: Sublagenicula brasiliensís, S. sinuate, Trileites tenais and Calomospora sp. The species of the genus Sublagenicula are dominant. Are here discussed the relationship presents among other species of megaspores too, and temporal and spacial distribution of that four species for Paraná Basin and Gondwana.
